Agonistic behaviour, made inspectable

Conflict Calculus

Before animals clash, they signal, sample, assess and sometimes withdraw. Enter a contest where the strongest-looking animal is not always the strongest, and a sensible decision can still produce an unlucky outcome.

Five species contexts Replayable cases AS91603 bridge No personal data

AI-generated natural-history illustration for species context. It is not a photograph, observation or data source.

ILLUSTRATIONBuilds species and habitat context. Never use it as data.
MODEL STATEShows a current hidden or visible variable in this teaching model.
MODEL OUTPUTResults generated from the named rules and replay code.
FIELD EVIDENCEA bounded claim linked to a real study and species.

01 · Contest lab

Read the signals before you call the clash.

Commit a prediction, reveal one cue at a time, then advise Contestant A as an observer. Your advice is recorded, not enacted. The seeded animals follow the disclosed teaching rule, allowing decision quality to be separated from a lucky or unlucky outcome.

Learning mode

Guided mode names the cue and provides a process prompt. Raw evidence remains visible in every mode.

Resource: breeding access

A roaring contest at dawn

Two adult stags meet near a harem. Both know their own condition better than they know their rival.

BEFORE STAGE 1
Anatomy is stylised and fixed across stages. Stage overlays and labelled readouts show the active model cue.
A cue hidden B cue hidden
Agonistic behaviourThe full contest repertoire, including display, threat, attack, submission, avoidance and withdrawal.
RHPResource-holding potential, a relative capacity to prevail. It is not the same as motivation.
Resource valueWhat the contested resource is worth to this individual in this context.
AssessmentUsing imperfect cues to update an estimate of the opponent or of one’s own capacity to persist.
EscalationMoving to a more costly or risky contest stage. It is not automatically the fitter choice.

02 · Matched cohort lab

One clash is a story. Repeated cases are evidence.

Change one factor, keep the random case stream matched, and compare 120 baseline contests with 120 treatment contests. Claims remain about this teaching model.

Design the comparison

Change one variable

LowerBaseline is 50Higher

Choose a directional prediction before running the cohort.

All settings are relative teaching-model indices. They are not field-calibrated probabilities or energy units.

MODEL OUTPUT · matched comparison

Baseline and treatment

Commit a prediction and run the paired cohort. A readable table will appear with the chart.

03 · Species transfer

Five biological contexts. One inspectable model.

Each context changes the resource, contest sequence, cue modality, reliability schedule and evidence boundary. All five currently use one shared teaching engine, so transfer is possible without pretending these are independently fitted species models.

1

Red deer

Acoustic and visual assessment. Roaring rate and vocal formants can carry different information, followed by parallel walking and possible antler contact.

Current mapping: staged cue reliability and a high-cost final stage inside the shared engine.

2

Fiddler crab

A large regenerated claw can resemble an original claw while producing lower force. The context makes imperfect honesty visible without implying conscious deception.

Current mapping: the shared engine plus a low-frequency regenerated-claw cue bias.

3

Stalk-eyed fly

Body size, eyespan, ownership and local context can covary with outcomes. The resource-value index represents context broadly, but female presence is not a separate control.

Current mapping: size cue, ownership and subjective value inside the shared engine.

4

Mantis shrimp

Meral displays may not settle size-matched contests. Ritualised strikes to an armoured telson can reveal persistence without ordinary lethal impact.

Current mapping: N. bredini stages and cue schedules inside the shared engine, not a fitted stomatopod model.

5

Hermit crab

Attackers rap a defender’s shell, and the shell’s value differs between individuals. Oxygen and stamina are important in the source biology but are represented only within composite RHP and cost, not as separate controls.

Current mapping: shell-value cue bias and cumulative stage costs inside the shared engine.

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Transfer question

Which visible pattern would distinguish assessment of an opponent from a contestant simply reaching its own cost threshold?

The answer needs experimental comparisons, not a dramatic animation.

04 · Learner guide

Follow the evidence chain.

The interface separates what exists in the case, what can be observed, what the model infers and what one outcome happens to show.

True state
Cue or display
Noisy estimate
Decision rule
Behaviour
Cost and outcome
Repeated payoff proxy
Screen elementTypeWhat it representsUse it to claim
Cinematic habitatILLUSTRATIONSpecies, resource and contest stage.Nothing quantitative.
Animal anatomy artworkILLUSTRATIONRecognisable morphology, fixed across contest stages and not cue-scaled.Nothing quantitative.
Signal overlays and readoutsMODEL STATEWhich signal is active and its seeded measured value.Which information was available in this model case.
Inference bars and ≈ ±1 SDMODEL OUTPUTA field observer’s teaching-model estimate and approximate propagated standard deviation.How uncertainty changed within the shared teaching rule.
Cohort distributionsMODEL OUTPUTRepeated results under matched seeds.A direction or difference inside this teaching model.
Species study noteFIELD EVIDENCEA bounded result from one named study.Only the species, setting and variable that study supports.
“Agonistic behaviour means aggression and fighting.”

No. It includes display, threat, submission, avoidance and withdrawal as well as physical aggression. A contest can be resolved without contact.

“The strongest animal should always win.”

Strength shifts the odds. Motivation, ownership, resource value, condition, information error and chance can all affect one outcome.

“The animal calculates the equation.”

No. The equation is our inspectable teaching representation of a conditional decision. It does not claim conscious arithmetic or purpose.

05 · Teacher guide

Teach the decision, not the spectacle.

The core move is from an appealing animal encounter to a controlled, repeated comparison and a bounded explanation.

Learning promise

Learners will distinguish agonistic behaviour from aggression, separate RHP from resource value, and use repeated model evidence to explain withdrawal or escalation under uncertainty.

Starting assumptions

Learners should know variation, competition, costs and benefits, and natural selection. They do not need prior knowledge of any exemplar species or probability equations.

Assessment use

Formative preparation for resource-based AS91603 reasoning. The tool does not grade the standard, establish a real animal mechanism or generate an assessment answer.

15 minutes · phenomenon
  1. Commit a winner prediction.
  2. Reveal one red deer contest.
  3. Separate decision quality from outcome.
  4. Exit with RHP versus value.
30 minutes · controlled test
  1. Pretrain five terms.
  2. Run a guided contest.
  3. Change one cohort variable.
  4. Write claim and evidence.
50 minutes · full inquiry
  1. Retrieve prerequisites.
  2. Predict and run.
  3. Complete two matched comparisons.
  4. Transfer to a second species.
  5. Write CER-U.
ElementDevelopingSecureStrong
ClaimNames an outcome.States a direction.States a bounded, conditional direction.
EvidenceUses one contest.Uses a cohort measure.Uses sample size, numerical contrast and matched design.
ReasoningRepeats “costs and benefits”.Links cue to decision.Links noisy assessment, value and future costs.
UncertaintyNo limit.Names model simplification.Names an alternative rule or field test.

Cognitive-science choices

  • Prediction before reveal creates generative effort.
  • Stages segment a complex process and keep the active question singular.
  • Labels sit beside the representation they explain.
  • Matched comparisons reduce confounding.
  • Immediate retrieval, delayed retrieval and species transfer revisit the causal model.
  • Scaffolds fade from Guided to Field Study to Exam.

Engagement without distortion

  • Authentic field-biologist decisions, not aggression points.
  • Surprising stochastic outcomes followed by explanation.
  • Replay codes support class comparison.
  • Progress is Predict, Control, Repeat, Explain and Critique.
  • No countdowns, streaks, leaderboards or rewards for escalating.
  • Accessibility and raw evidence never sit behind an achievement gate.

Welfare boundary: learners must not provoke, stage or approach real animal contests. Use published observations, remote footage or ethically collected datasets.

06 · Behind the model

An inspectable decision under uncertainty.

The mathematics is a teaching model, not a claim that animals perform arithmetic. Species evidence anchors cue direction and contest structure. Classroom weights remain disclosed assumptions.

True stateRHP, condition, ownership, resource value
ObservedSpecies cue plus seeded measurement noise
EstimatedUpdated A and B values plus a displayed model standard deviation
DecisionExpected resource gain compared with future cost
EvidenceStage log, paired cohort, raw summary table
field estimatet = reliability × observed cue + (1 − reliability) × previous estimate
starting teaching prior = 50 ± 25
cue-noise SD = 23 × (1 − reliability) + 3
model uncertainty SDt ≈ √{[(1 − reliability) × previous SD]² + [reliability × cue-noise SD]²}
Pmodel(A wins) = logistic[slope × (estimate A − estimate B)]; Pmodel(B wins) = 1 − Pmodel(A wins)
continue value for A = P(win) × resource value − expected display, energy and injury costs
model recommendation = withdraw if continue value < 0; otherwise sample or escalate according to stage
Presolved fight(A wins) = logistic[slope × (true RHP A − true RHP B)]
long-run payoff proxy = resource gains − display costs − injury costs across repeated contests

Implementation disclosure: every species context currently uses this same decision engine. Species selection changes the staged behaviour, cue labels, stage reliability, default resource and injury indices, logistic slope, and two bounded cue modifiers: regenerated-claw unreliability for fiddler crabs and shell-value bias for hermit crabs. The uncertainty propagation assumes independent stage-cue errors; clipping at 0 and 100 makes the displayed SD approximate. These settings are teaching assumptions, not fitted population parameters.

What is empirical

  • The species and documented contest behaviours.
  • The direction of selected cue, ownership, resource and stamina relationships.
  • The existence of imperfect or dishonest signals in named systems.
  • The distinction among mutual, self and cumulative assessment hypotheses.

What is a teaching assumption

  • The 0 to 100 indices and exact thresholds.
  • The normal cue-noise distribution and logistic win curve.
  • One shared decision engine across five species contexts.
  • The classroom injury proxy and payoff scale.
  • Any claim that this rule is the uniquely correct mechanism.
PresetField evidenceTeaching assumptionDo not concludePossible field test
Red deerRoaring rate and formants can inform rivals; visual escalation follows.Three cues update one shared estimate.Pitch alone reveals strength, or all stags use one rule.Playback of controlled formants and rate with matched rivals.
Fiddler crabRegenerated claws can overstate force while resembling original claws.A small seeded share of cases carries a bluff penalty.Crabs consciously lie, or every large claw is reliable.Compare initiation and escalated outcomes by claw type and residency.
Stalk-eyed flyBody size, eyespan and ownership covary with outcomes; female presence can alter local value.The shared engine maps alignment to a noisy body-size estimate; female presence is not a separate state variable.Relative eyespan is an independent strength gauge or all diopsids match.Factorial manipulation of body size, ownership and lek value.
Mantis shrimpSize-matched contests often escalate to repeated telson strikes; evidence supports mutual assessment in tested N. bredini.Stage cues update opponent estimates and persistence shifts withdrawal.Meral spread reliably predicts strike force, or all stomatopods share this rule.Separate size mismatch, display, strike number, role and residency.
Hermit crabShell rapping and oxygen availability affect attacker persistence.The shared engine maps performance cues and shell value to a threshold; oxygen is omitted as a separate variable.Duration alone proves cumulative or mutual assessment.Manipulate oxygen, shell value and opponent action independently.

07 · Product and marketing lens

What the experience is selling.

The strongest proposition is not “watch animals fight”. It is “learn to make a defensible decision from incomplete biological evidence, then test it across cases”.

Distinctive hook

Decision quality is separated from luck. Learners can make a biologically sensible recommendation and still see an upset, which creates a natural reason to run repeated cases.

Visual signature

Cinematic habitat, fixed stylised anatomy, stage-specific signalling, a calculated model uncertainty measure and a forensic reveal. The interface moves from atmosphere to evidence without confusing the two.

Classroom value

One case captures attention. Matched cohorts support graph and table interpretation. Cross-species transfer prevents memorising a deer-only story.

Safe public claim

“Explore how animal contests can be resolved through signals, assessment, withdrawal and escalation, then test explanations against replayable data.”

Avoid claiming

Species-accurate prediction, validated learning gains, automatic NCEA grading, a universal contest equation or a proven mechanism in every species.

Pilot question

After one 50-minute lesson, can learners distinguish RHP, resource value and outcome luck, then transfer the causal model to an unfamiliar species dataset?

GraphicWhat changes itWhat it communicatesMarketing roleEvidence status
Habitat artworkSpecies selectionOrganism, scale, resource and moodImmediate curiosity and recognisable identityIllustration only
Signal overlay and readoutContest stage and seeded cueInformation is released over time, while anatomy remains fixedThe signature “read the rival” interactionModel state
Inference barsRevealed noisy cuesEvidence changes confidence without certaintyVisible intellectual progressModel output
Decision/outcome splitUser recommendation and stochastic resultSensible decisions can have unlucky outcomesMemorable learning twistModel output
Matched cohort barsOne controlled variableDistribution-level evidence and trade-offsTeacher credibility and exam preparationModel output
Study ledgerActive species source packEmpirical, simplified, omitted and uncertainTrust and scientific transparencyField evidence plus disclosure

Recommended build order

  1. Pilot the shared red deer contest and cohort engine.
  2. Validate the fiddler-crab regenerated-claw cue modifier.
  3. Test Exam mode and the teacher route.
  4. Only then prototype genuinely different assessment engines for mantis shrimp and hermit crabs.
  5. Use the stalk-eyed fly as a transfer case.
  6. Pilot before a separate population-genetic Evolution Lab.

Media and rights

The two raster artworks in this prototype were generated for this project and are explicitly labelled as illustration. Production should retain prompts and provenance. Any future field photograph, audio, video or spectrogram needs file-level creator, licence, crop and modification records.

Do not hotlink images or imply that generated art depicts a measured trial. For taonga species, seek suitable authority, provenance and permission before use.

Return to the contest

08 · Source ledger

Primary science and learning design.

Sources anchor directions and boundaries. The simulator remains an explicit teaching model unless and until parameters are fitted and independently validated for a defined field population.

Curriculum

NZQA Standard 91603 and the 2026 Level 3 Biology assessment specification, which names agonistic behaviour and graphical and tabulated data.

Fiddler crab

Backwell et al., 2000; Morrell et al., cumulative-assessment analysis; Lailvaux, Reaney and Backwell, 2009. Sources use the former genus name Uca; this prototype uses Austruca mjoebergi.

Mantis shrimp

Green and Patek, 2015, telson sparring; Green and Patek, mutual-assessment analysis. A 2026 study of related Gonodactylus childi found role and individual variation, so the rule is not generalised across mantis shrimp.

Hermit crab

Briffa and Elwood, 2000, oxygen availability and alternative assessment rules in shell fights.

Feedback and guidance

Shute, 2008; Kalyuga et al., 2003.